We have a pretty decent line-of-descent for various yersiniae. It started as something you'd get in the wild, but not spread (much) person-to-person. It was, basically, like anthrax. Speaking-of, the bug has been found in a sheep. This one still lacks the Gained Function (as it were) which made it a flea bug, which they're calling for 1800s BC.
Which calls into question the sorts of yersinia as afflicted Cucuteni - and BellBeaker Somerset. These villages, unlike Eurasia, had granaries and urban rodents. Did their fleas not carry le Peste? Perhaps these first rodents were spreading the bug like a hanta.
After getting those mutations which made yersinia flea-transmissible, and once it got into rat fleas, humanity was over.
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