The "R" Y-chromosome featured in Mal'ta Boy, in Siberia. R1 is ancestor to Indo-Europeans... and to North Americans. Already at the beginning of our Holocene, R1a had split off in the Russias. I sometimes slip up and consider R1b, R2b. So: how about those other R's?
R2 clusters in Central Asia and Pakistan. These regions also have the miscellaneous R*, Mal'ta aside. Also during ice ages I expect populations to stick with the warmer, more-sheltered regions.
At a guess, Mal'ta was peripheral to R's Pleistocene homeland. That homeland was the greater northern Iran between the Caspian and the Aral. The domestication of the dog and [UPDATE 1/25/21: then] Solutrean technology (from the Catalan coast) allowed R1 to expand permanently into the Baltic (R1a), the Danube (R1b) and Siberia (R1-Bering). Meanwhile R2 and the rest stayed home.
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