Among the polities the Inca took over were the Chincha. The reason the Inca wanted that site was guano, which is not a Spanish word; here from seabird.
The paper claims the Moche already knew of this resource, which fertilised - what else - maize. Around AD 1250 the Chincha start using bird imagery on their pots. Possibly the birds' pots; DNA analysis might pick it up.
I do not find noted in this paper, the Huari. Perhaps because the Chincha were too far south, better contemporaries with the Tiahuanaco.
Around AD 1400, the Inca came knocking and annexed this kingdom. This is about when the Inca spread fame of the "Lunahuaná", that is the guano people - north of the Chincha. The paper reports that the Spaniards will observe that the Chincha lord ranked among the high notables in the Inca retinue.
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